Puffinus carneipes; Print; The flesh-footed shearwater (Ardenna carneipes; formerly Puffinus carneipes) is a medium-sized shearwater. Its plumage is black. It has pale pinkish feet; and a pale bill with a distinct black tip. Together with the equally light-billed pink-footed shearwater; it forms the Hemipuffinus group; a superspecies which may or may not have an Atlantic relative in the great shearwater. These large shearwaters are among those that have been separated into the genus Ardenna. Recent genetic analysis indicates evidence of strong divergence between Pacific colonies relative to those in South and Western Australia; thought to be explained by philopatry and differences in foraging strategies during the breeding season.; 1820-1860